1. Key Laboratory of Stress Plant Research, College of Life Sciences, Shandong Normal University, Jinan 250014, China;Shandong Shengli Co., LTD, Jinan 250101, China 2. Key Laboratory of Stress Plant Research, College of Life Sciences, Shandong Normal University, Jinan 250014, China 3. Key Laboratory of Stress Plant Research, College of Life Sciences, Shandong Normal University, Jinan 250014, China;Weifang Medical University, Weifang 261042, China 4. College of Life Sciences, Qufu Normal University, Qufu 273165, China 5. Jinan No. 3 Vocational School, Jinan 250001, China