1(State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, the Chinese Academy of Sciences, Beijing 100093, China) 2(Biological Institute, Showa Pharmaceutical University, 3-Chime, Higashi-Tamagawagakuen, Machida, Tokyo 194-8543, Japan) 3(College of Forestry, the Northeast Forestry University, Harbin 150040, China) 4(Department of Botany, United States National Herbarium, National Museum of Natural History, MRC-166, Smithsonian Institution, Washington, DC 20013-7012, USA)
