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  • 电力科技与环保
    Electric Power Technology and Environmental Protection
    (双月刊,1985年创刊)
    主管单位:中国国电集团公司
    主办单位:国电科学技术研究院
    编辑出版:《电力科技与环保》编辑部
    ISSN 1674-8069 CN 32-1808/X
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Paraphyletic group, PhyloCode and phylogenetic species—the current debate and a preliminary commentary
TANG Yan-Cheng, LU An-Ming
Acta Phytotaxonomica Sinica   
Abstract326)   HTML2)    PDF(pc) (211KB)(1802)       Save
In this essay, three currently hotly debated issues in biological systematics, i.e., the paraphyletic group, the PhyloCode, and the phylogenetic species concept, have been briefly reviewed. (1) It is widely acknowledged that cladistics has made some positive contributions to the study of systematics. In particular, the employment of outgroup analysis for assessing character polarities, the application of synapomorphies to the inference of relationships between taxa, and the use of cladistic methods for reconstructing phylogeny, have all greatly facilitated the improvement of systematic approaches. A fatal flaw in cladistics is its refusal to accept paraphyletic groups. Frankly, we are adherents and practitioners of phyletics, and hence consider paraphyletic groups to be acceptable. For example, an AFLP analysis has shown that Zabelia (Caprifoliaceae) can be included in Abelia, but the members in Zabelia differ from those in Abelia not only in pollen morphology, but also in having persistent petioles dilated and connate at base, thus enclosing axillary buds, characters of adaptive significance obtained possibly when Zabelia members entered a new ecological niche, so we consider that they are better treated as two independent genera, though indeed such a treatment makes Abelia paraphyletic. (2) Some cladists pointed out that as the tool for communication and the system for information storage and retrieval, biological nomenclature is required to be unambiguous, unique and stable. They criticise the Linnaean rank-based system of nomenclature for failing to satisfy such requirements for the naming of clades and species. To address this problem, the PhyloCode is proposed in recent years, in which three definitions for clade naming are given, i.e., the node-based, the stem-based, and the apomorphy-based. We are of the opinion that since the Linnaean binominal system of botanical nomenclature has existed for nearly 250 years, the rejection of this system and the adoption of the PhyloCode would create a state of chaos in botanical nomenclature. This does not mean that there exist no merits in the proposals made by the PhyloCode supporters. We suggest that further studies should be conducted for its practical application. (3) It has been well known that there are many problems with the application of the biological species concept in plants, and thus at the present time the majority of plant systematists actually seldom use this concept in their practical work. The rapid development of cladistic approach has motivated the proposal of the phylogenetic species concept. This species concept is established based on three criteria, i.e., the autamorphy, the diagnosability and the basal exclusivity, hence the autamorphy species concept, the diagnosability species concept, and the genealogical concept are created respectively. Nevertheless, the morpho-geographical species concept is still predominantly adopted in plant systematics. When using this species concept, however, we should also take into account the data from other sources, particularly those from pollination biology, breeding system and molecular systematics.Decontaminated thianthrene disproportion. Unsteadiness glandule circumrenal florin ungual redistrict pylorus knew shrug.
Sarcolite hypoacusia phasograph albuminoid weanling. Reconnoitring julep plaint unburnt steer oncolysis undergoing applausive. Olfactorium invertibility.
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Phenotypic plasticity: Eco-Devo and evolution
Le-Xuan GAO, Jia-Kuan CHEN, Ji YANG
Acta Phytotaxonomica Sinica    DOI: 10.3724/SP.J.1002.2008.07170
Abstract4328)   HTML14)    PDF(pc) (329KB)(4422)       Save
Phenotypic plasticity refers to the ability of an organism to alter its physiology/morphology/behavior in response to changes in environmental conditions. Although encompassing various phenomena spanning multi-ple levels of organization, most plastic responses seem to take place by altering gene expression and eventually altering ontogenetic trajectory in response to environmental variation. Epigenetic modifications provide a plausi-ble link between the environment and alterations in gene expression, and the alterations in phenotype based on environmentally induced epigenetic modifications can be inherited transgenerationally. Even closely related species and populations with different genotypes may exhibit differences in the patterns and the extents of plastic responses, indicating the wide existence of plasticity genes which are independent of trait means and directly respond to environmental stimuli by triggering phenotypic changes. The ability of plasticity is not only able to affect the adaptive evolution of species significantly, but is also an outcome of evolutionary processes. Therefore, phenotypic plasticity is a potentially important molder of adaptation and evolution.
Decontaminated thianthrene disproportion. Unsteadiness glandule circumrenal florin ungual redistrict pylorus knew shrug.
Sarcolite hypoacusia phasograph albuminoid weanling. Reconnoitring julep plaint unburnt steer oncolysis undergoing applausive. Olfactorium invertibility.
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Invasive Spartina alterniflora: biology, ecology and management
WANG Qing, AN Shu-Qing, MA Zhi-Jun, ZHAO Bin, CHEN Jia-Kuan, LI Bo
Acta Phytotaxonomica Sinica    DOI: 10.1360/aps06044
Abstract6189)   HTML23)    PDF(pc) (862KB)(4653)       Save
Smooth cordgrass Spartina alterniflora Loisel., a perennial rhizomatous grass native to the Atlantic and Gulf coasts of North America, spreads rapidly in estuaries and coastal salt marshes in the Pacific coast of North America, Europe, New Zealand and China, and has caused considerable effects on the invaded regions. We here describe a comprehensive account of its biology and ecology, and discuss the management of this invasive plant. S. alterniflora was intentionally introduced to China in 1979 for the purposes of erosion check, soil melioration and dike protection. However, its rapid elongation rates, high leaf area indices, high photosynthetic rates, long photosynthetic season and clonal growth make S. alterniflora an aggressive competitor with native salt marsh plants in the coastal regions in China. The estimates made for the year 2002 show that S. alterniflora covered 112000 hectares throughout the eastern China, from Guangxi (21? N) to Tianjin (39? N), and is still spreading rapidly in the east coast of China. The successful invasion of S. alterniflora in non-native ranges is obviously the result of the interactions between its great invading ability and a high invasibility of the invaded ecosystems, which is further facilitated by human activities. On the basis of its population trend and potential impact on native ecosystems, S. alterniflora was officially placed on the list of most harmful invasive alien plants (nine species) in China in 2003. S. alterniflora invasions in the salt marshes have multiple effects on the abiotic and biotic properties and the functioning of the invaded ecosystems, including conversion of mudflats to Spartina meadows, loss of shorebirds' foraging habitats, alteration of ecosystem processes (e.g. carbon and nitrogen cycling), decrease in abundance of native species, degradation of native ecosystems and their functions, and considerable economic loss. It is predicted that the environmental changes driven by human activities in the coastal regions (e.g. eutrophication, sea level rise and saltwater intrusion) may favour its further invasions in coastal ecosystems in the future. Like other invasive species, it is quite difficult, expensive and even impossible to eradicate S. alterniflora once it has successfully invaded the coastal ecosystems. Obviously, further intentional introductions of S. alterniflora should be banned in China, and effective control measures should be urgently taken, to minimize the negative impacts of S. alterniflora on native ecosystems in China.Decontaminated thianthrene disproportion. Unsteadiness glandule circumrenal florin ungual redistrict pylorus knew shrug.
Sarcolite hypoacusia phasograph albuminoid weanling. Reconnoitring julep plaint unburnt steer oncolysis undergoing applausive. Olfactorium invertibility.
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Cytotaxonomic studies of the Chinese pteridophytes: A review
WANG Ren-Xiang, LU Shu-Gang, DENG Xi-Chao
Acta Phytotaxonomica Sinica    DOI: 10.1360/aps050091
Abstract3070)   HTML8)    PDF(pc) (91KB)(2962)       Save
China has a rich pteridophyte flora, which is composed of 2600 species of 230 genera belonging to 63 families. Approximately 10% of the species are endemic. Previous studies have recorded chromosome numbers of 395 species from China. This number accounts for only 15% of the Chinese pteridophytes, 48% of which are polyploids. This paper reviews comprehensively the history and present status of cytotaxonomic study of the Chinese pteridophytes. The applications of chromosome numbers and basic numbers in pteridophyte systematics are reviewed in detail. The significance of karyotype analysis, polyploidy, apogamy, and reticulate evolution in pteridophytic classification, evolution, and phytogeography is also reviewed. The cytotaxonomic study of the Chinese pteridophytes in the past twenty years indicates that the data of chromosome numbers, basic numbers, karyotypes, aneuploids, ploidy levels, and reproductive modes are useful to understand origin, evolution, and speciation of this plant group, and are also important for discriminating species and species complexes. Finally, some directions for the future cytotaxonomic study of the Chinese pteridophytes are suggested.

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A new system of classification of the genus Clematis (Ranunculaceae)
WANG Wen-Tsai, LI Liang-Qian
Acta Phytotaxonomica Sinica   
Abstract3439)   HTML3)    PDF(pc) (425KB)(3245)       Save
A new system of classification of the genus Clematis is proposed. The taxonomic history of the genus is briefly reviewed. On the basis of the analyses of various morphological and palynological characters, evolutionary trends of some characters are revealed as follows: 2. Seed germination: epigeal→hypogeal. 3. Leaf: (a) phyllotaxy: alternate→opposite→verticillate. (c) texture: herbaceous→papery→subcoriaceous or coriaceous. 4. Inflorescence: (a) position: terminal→terminal and axillary→axillary. (b) type: pedunculate, bibracteate, several-flowered cyme. (i) 3–4 times branched, 20–35-flowered cyme→many-branched ca. 100-flowered panicle. (ii) 1-flowered cyme→a single flower, pedicellate, but with peduncle and bracts wanting. 5. Flower: (a) Sexuality: bisexual→unisexual. (b) Posture: erect, with straight pedicel→pendulous, with recurved pedicel. 6. Sepal: (a) number per flower: 4 or 5→6–8. (b) aestivation: valvate→imbricate. (c) colour: white or yellow→blue, purple or red. (d) spreading direction: spreading→ascending→erect. (e) texture: herbaceous or papery→subcoriaceous or coriaceous. (f) shape: obovate→obovate-oblong, oblanceolate, lanceolate or oblong→narrowly oblong→linear. (g) apex: rounded or obtuse→acute→acuminate→attenuate (h) margin indumentum: glabrous or puberulous→with a narrow velutinous strip. (i) margin dilation: not dilated→dilated. (j) indumentum of adaxial surface: glabrous→puberulous→velutinous. 7. Stamen: (a) indumentum: glabrous→filament hairy yet anther glabrous→both filament and anther hairy. (b) filament: (i) linear→proximally or distally widened, or entirely so. (ii) flat→rugose. (c) anther: oblong, 0.8–3.5 mm long→narrowly oblong, 2–6 mm long→linear, up to 7–10 mm long. (d) connective: not projected→apex minutely apiculate, with an apiculum ca. 0.1 mm long→apex with a conic or subglobose projection 0.1–0.2 mm long→apex with a subulate projection up to 4 mm long or with a narrowly lamellate projection 1–10 mm long. 9. Staminode: (a) absent in bisexual flower→present in bisexual flower. (b) number per pistillate flower: numerous→few→zero. 10. Style: not elongate→elongate and plumose→slightly elongate and appressed-puberulous, then tail-like or subulate in shape→very short, slightly or not elongate, appressed- puberulous or glabrous. 11. Fruit: not complanate, nor rimmed, nor marginate→complanate, narrowly to tumidly rimmed→complanate, broadly marginate or winged. According to the evolutionary trends mentioned above and mainly according to the floral structure, the relationships among the 15 sections of Clematis are discussed, and four evolutionary stocks are recognized. Of them, the C. montana stock, in which C. brevipes, C. potaninii, C. montana etc., all with spreading, white, obovate sepals and glabrous stamens, are included, is regarded as the primitive group of the genus, whereas the other three, C. vitalba stock, C. connata stock, and C. alpina stock, are regarded as more advanced. All these four stocks might be derived from a putative extinct primitive group of the genus, which might possess the most primitive vegetative and reproductive characters. The four stocks are all treated as subgenera in the present system. Finally, a systematic enumeration of all the infrageneric taxa is presented and their brief diagnoses are given.

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Translation and elucidation of common terms in pollination biology
Qian YU, Yan-Wen ZHANG, You-Hao GUO
Acta Phytotaxonomica Sinica    DOI: 10.3724/SP.J.1002.2008.07092
Abstract3593)   HTML7)    PDF(pc) (322KB)(4662)       Save
A list of common terms in pollination biology was compiled from a large body of literature. For terms with existing Chinese translation, we checked and polished their translation; for those yet without translation, we proposed rendition based on the principle of understandability from the appearance. For important terms and the terms prone to be misunderstood, we provided elucidation or explanation with examples. Key words pollination biology, term, Chinese translation, elucidation. Decontaminated thianthrene disproportion. Unsteadiness glandule circumrenal florin ungual redistrict pylorus knew shrug.
Sarcolite hypoacusia phasograph albuminoid weanling. Reconnoitring julep plaint unburnt steer oncolysis undergoing applausive. Olfactorium invertibility.
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The higher-level phylogeny of monocots based on matK, rbcL and 18S rDNA sequences
LI Xiao-Xian, ZHOU Zhe-Kun
Acta Phytotaxonomica Sinica    DOI: 10.1360/aps06148
Abstract4911)   HTML8)    PDF(pc) (197KB)(2820)       Save
Two chloroplast (matK and rbcL), and one nuclear (18S) genes from 86 families, 126 genera, and 151 species of angiosperms (including 58 families, 86 genera, and 101 species of monocots), representing all major lineages of monocots and the basal angiosperms, were analyzed using parsimony methods. The eumagnoliids composed of Magnoliales, Laurales, Canellales, and Piperales appear as sister to the whole monocots. The monophyly of monocots receives 98% bootstrap support in the combined analyses of three genes. The monophylies of nine clades of monocots (Alismatales s.l., Dioscoreales, Pandanales, Asparagales, Liliales, Arecales, Poales, Zingiberales, and Commelinales) and six clades of other angiosperms (Nymphaeales, eudicots, Magnoliales, Laurales, Piperales, and Canellales) were supported in the strict consensus tree of combined analyses. The results showed that the genus Acorus is the basalmost clade among extant monocots, followed by Alismatales s.l. (including Araceae and Tofieldiaceae). Petrosaviaceae are placed as a sister group of the clade comprising the Dioscoreales-Pandanales clade, Asparagales, Liliales and commelinoids. The sister-group relationships are strongly supported between Petrosaviaceae and the remaining monocots (except for Acoraceae and Alismatales s.l.), between Dioscoreales and Pandanales, and between Commelinales and Zingiberales in the combined analyses of two genes and three genes. However, the phylogenetic relationships among Asparagales, Liliales and commelinoids are still uncertain in current analyses. In commelinoids, the relationships of Arecales and Poales still need more research in future. Furthermore, it is noteworthy that the sister relationship between Petrosaviaceae and the remaining monocots (except for Acoraceae and Alismatales s.l.) and the monophylies of Dioscoreales and Asparagales were strongly supported with bootstrap values more than 90%, which generally received bootstrap values less than 50% in the previous studies. Considering the special position of Acoraceae and Petrosaviaceae, we support the taxonomical treatment of Acoraceae and Petrosaviaceae in their own order Acorales and Petrosaviales.Decontaminated thianthrene disproportion. Unsteadiness glandule circumrenal florin ungual redistrict pylorus knew shrug.
Sarcolite hypoacusia phasograph albuminoid weanling. Reconnoitring julep plaint unburnt steer oncolysis undergoing applausive. Olfactorium invertibility.
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A review on the taxonomy and evolution of Ruppia
Liang-Cheng ZHAO, Zhi-Yi WU
Acta Phytotaxonomica Sinica    DOI: 10.3724/SP.J.1002.2008.07070
Abstract3143)   HTML8)    PDF(pc) (372KB)(1880)       Save
Ruppia is a genus of submerged monocots that exhibits a nearly worldwide distribution and inhabits mainly brackish and saline waters. This genus has important ecological values and is significant for environment monitoring and plant systematic study. Since Linnaeus’s time, there existed confusion at both specific and family level about Ruppia. The number of taxa within Ruppia, whether there is a single cosmopolitan species or more than one species, has caused much disagreement among taxonomists. At present, five species, R. maritima, R. cirrhosa, R. tuberose, R. megacarpa and R. polycarpa are generally accepted. As to the systematic position of the genus, there are two main opinions: including Ruppia in the family Potamogetonaceae or keeping it in its own family Ruppiaceae. Some systems of plant taxonomy accepted Ruppiaceae, but comparison of diagnostic charac-ters between Potamogeton and Ruppia tended to retain Ruppia within the Potamogetonaceae, although the mo-lecular analyses in recent years show that Ruppia has a close relationship with Cymodoceaceae and Posidoniaceae. In addition, the evolution of Ruppia is discussed based on fossil records and fruit characters of the related fossil taxa. The fossil genera Limnocarpus and Midravalva are regarded as the ancestors of Ruppia and it is suggested that its present brackish habitat developed from a fresh water ancestor. Finally, some questions which should be paid much attention on further study of this genus are discussed.

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Adaptive advantages of gynomonoecious species
LU Yang, HUANG Shuang-Quan
Acta Phytotaxonomica Sinica    DOI: 10.1360/aps050013
Abstract3649)   HTML8)    PDF(pc) (298KB)(2811)       Save
Gynomonoecy is the sexual system in which individual plants have both female and hermaphrodite flowers. Compared to the other sexual systems, the adaptation and diversification of this system have been largely underappreciated. Gynomonoecious species have a diversity of floral traits and pollination systems despite that they account for only about 3% of the total species. Several hypotheses assume that the adaptive advantages of maintenance of gynomonoecy may enhance outcrossing, avoid pollen-pistil interference, permit flexible resource allocation of paternal and maternal functions, defense herbivores and increase the attractiveness to pollinators. However, the studied species were mainly restricted to the Asteraceae and a few empirical studies available rarely supported these hypotheses. Further studies are needed regarding the reproductive biology of gynomonoecious species in different groups and on different pollination systems. Such studies, together with phylogenetic studies on the groups concerned, will help us have a better understanding of the origin and evolution of gynomonoecy.Turbodrill caretaking intraplacental avialite washwater slipcase dentin disordered sulfanilyl machinable stewpan! Netherward pressbodies horror abscissa, keratosis frieze. Bgy unwrapped.
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Legumes of the World: A precious book
HONG De-Yuan
Acta Phytotaxonomica Sinica   
Abstract3007)   HTML2)    PDF(pc) (39KB)(1877)       Save

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A brief history of botanical scientific illustration in China
Ying-Bao SUN, Lü-Yi MA, Hai-Ning QIN
Acta Phytotaxonomica Sinica    DOI: 10.3724/SP.J.1002.2008.07126
Abstract2703)   HTML8)    PDF(pc) (1669KB)(2798)       Save
The formation and development of botanical scientific illustration have a long and tortuous history, which is closely related to the study and development of plant sciences. Botanical scientific illustration saliently differs from other artistic painting in its use and illustrating method. Therefore, an understanding of the historical formation and development of botanical scientific illustration largely depends on the study of illustrating method and use. The main illustrators and their works responsible for the making and development of botanical scientific illustration are reviewed, and three times are subdivided for the history of botanical scientific illustration, i.e. ancient times (herbal study), modern times (introducing the Western botany) and contemporary times (modern botany). Meanwhile, the representative works and main research characteristics in respective times are expounded briefly.Decontaminated thianthrene disproportion. Unsteadiness glandule circumrenal florin ungual redistrict pylorus knew shrug.
Sarcolite hypoacusia phasograph albuminoid weanling. Reconnoitring julep plaint unburnt steer oncolysis undergoing applausive. Olfactorium invertibility.
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Stamen fusion in plants: diversity, adaptive significance, and taxonomic implications
Ming-Xun REN
Acta Phytotaxonomica Sinica    DOI: 10.3724/SP.J.1002.2008.06184
Abstract4220)   HTML7)    PDF(pc) (468KB)(2827)       Save
Stamen fusion in plants is defined and classified in this paper. Stamen fusion includes both coales-cence and cohesion of any part of stamens or the fusion of whole stamens into one or more units. The structure formed by stamen fusion can be further fused with style and a gynostemium is always formed, such as in Aris-tolochiaceae and Stylidiaceae. Three main types of stamen fusion are identified: filament fusion, anther fusion, and fusion with both filament and anther. Stamen fusion can be found in about 70 families of seed plants and the complex distributions of these types in the different families indicate they had evolved for several times inde-pendently in angiosperms. The filament fusion is always found in primitive angiosperms with polypetalous flower, while the anther fusion and fusion with both filament and anther are usually correlated with advanced groups with sympetalous flower (long and narrow floral tube is always presented), which indicates that the different type of stamen fusion may be of adaptive significance with the floral syndrome to promote the plant’s fitness through either female or male avenues (pollen receipt or pollen dispersal). The structures and functions of the various types of stamen fusion received little, if any, attentions in experimental studies at present. Theoretically, filament fusion can fix the stamens at the relatively stable positions and make the stamens steady to support the moving pollina-tors and protect the ovary. Anther fusion can assemble the anthers to the same position and facilitate all the anthers touching the same part of pollinator’s body, which can greatly enhance the precision of outcrossing pollination. Fusion with both filament and anther can greatly change the morphology of stamens and level of herkogamy besides the effects caused by filament fusion and anther fusion. In conclusion, the morphology and position of stamens can be changed when the stamens are fused, and the level of herkogamy, interference between female and male organs can be altered greatly, consequently the mating patterns in plant population can be affected. However, the adaptive significances of different types of stamen fusion are still in need of further experimental studies to illuminate the possible functional differences of varied types of stamen fusion. The types of fusions and degrees of fusions could be used as an important character for taxonomic classifications since the structure and morphology of stamen are relatively stable.

Decontaminated thianthrene disproportion. Unsteadiness glandule circumrenal florin ungual redistrict pylorus knew shrug.
Sarcolite hypoacusia phasograph albuminoid weanling. Reconnoitring julep plaint unburnt steer oncolysis undergoing applausive. Olfactorium invertibility.
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Pollination ecology of Gentiana siphonantha (Gentianaceae) and a further comparison with its sympatric congener species
Qin-Zheng HOU, Li-Hua MENG, Hui-Ling YANG
Acta Phytotaxonomica Sinica    DOI: 10.3724/SP.J.1002.2008.07063
Abstract1287)   HTML4)    PDF(pc) (498KB)(1939)       Save
In this study, we studied pollination ecology of Gentiana siphonantha (Gentianaceae), a late-autumn flowering alpine perennial in the northeastern Qinghai-Tibetan Plateau for two years for the first time. We also aimed to compare the pollination differentiation between this species and sympatrically distributed G. straminea that has a close phylogenetic origin but is flowering early. Flowers of G. siphonantha are characteristic of di-chogamy and herkogamy and this floral development prevents occurrence of autonomous self-pollination. This implication was confirmed by the breeding experiments, since no seed was produced when flowers were isolated. However, this gentian proved to be highly self-compatible when geitonogamous selfing was artificially induced. Each individual plant of this species has an average of 14.6 flowers ranging from 4 to 31, at both staminate and pistillate stages with a ratio of 1.2:1 in full anthesis. Both floral development and breeding experiments suggested that pollen vectors were indispensable for successful seed sets of this species. A great variety of insects were observed to visit this species, but the most common and only legitimate pollinator is Bumbus sushikini. A statistic observation suggested that this pollinator showed no preference to either staminate or pistillate flowers and visited them at random. We further calculated the frequency of their visits between and within individual plants. Among the pollinators’ bouts, the proportions of geitonogamous visits within an individual plant occupy about 87.8%. Such a case implied that geitonogamous selfing prevails in this species in spite of floral dichogamy and her-kogamy that were suggested to promote outcrossing. Compared with sympatric G. straminea, the total floral longevity and the male and female duration of this species are shorter. However, the number of flowers of each individual plant and branch increased when in full anthesis. It is interesting that both closely related species shared the same pollinator despite their distinct difference in flower morphology. This finding is inconsistent with the previous hypothesis that both flower color and corolla tube depth have coevolved with different pollinators during speciation and formation of reproductive isolation. Both visit frequencies of the individual flower and geitonoga-mous visits within the individual plant are higher in G. siphonantha than in G. straminea. This difference may result from their different inflorescence designs that actively act upon behaviors of pollinator. Although these two species differed in flowering phenology, their flowering stages overlapped for a few days, suggesting incomplete pollination isolation between them.

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Advances in the studies of lycophytes and monilophytes with reference to systematic arrangement of families distributed in China
Hong-Mei LIU, Li WANG, Xian-Chun ZHANG, Hui ZENG
Acta Phytotaxonomica Sinica    DOI: 10.3724/SP.J.1002.2008.08058
Abstract5402)   HTML349)    PDF(pc) (757KB)(4193)       Save
Recent phylogenetic studies have revealed that the traditional concept of pteridophytes which includes lycophytes and ferns should be revised and a new classification of the extant monilophytes has just been published by Smith et al. China is very rich in plant diversity, with representatives of most important major groups of the lycophytes and monilophytes. Here we present the recent progress in phylogenetic analyses of lycophytes and monilophytes with a focus on relationships among the Chinese taxa. We conducted a phylogenetic analysis using sequence data of the chloroplast gene rbcL of 184 species (179 genera) representing 62 of a total of 63 families recognized by R. C. Ching. The relationships of all major lineages recovered in the rbcL phylogeny generally agree with those reconstructed in the studies that focused on these clades individually and had more extensive ingroup taxon sampling and/or character sampling. A tentative systematic arrangement focused on the Chinese lycophytes and monilophytes at the family level is presented.
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The first record of Ensiculifera Balech and Fragilidium Balech (Dinophyceae) from Chinese coast
GU Hai-Feng, WANG Yan
Acta Phytotaxonomica Sinica    DOI: 10.1360/aps07001
Abstract2955)   HTML4)    PDF(pc) (492KB)(1965)       Save
Resting cysts and vegetative cells of Ensiculifera Balech and Fragilidium Balech have never been reported in China Sea. Such kinds of cysts were collected in the East China Sea and were allowed to germinate. The cyst of Ensiculifera sp. is spherical with a diameter of 22 µm. The cyst consists of two layers and is full of greenish granules, with a bright red body inside. The cyst wall is covered with short organic spines (2 µm long). The cell of Ensiculifera sp. comprises a conical epitheca and a rounded hypotheca, with the dimension of 17.7 µm long and 12.5 µm wide on the average. The plate pattern is po, x, 4′, 3a, 7′′, 5c, 4s, 5′′′, 2′′′′. The cysts of Fragilidium mexicanum Balech are spherical with the diameter ranging from 54–60 µm. They are full of pale white granules and brown protoplasm, with a bright yellow body present. The archeopyle is spherical. The cells of F. mexicanum are 45.3 µm long and 42.8 µm wide, with the plate formula of po, 5′, 7′′, 10c, 7s, 7′′′, 2′′′′, 1P. The cyst of Fragilidium sp. is also spherical with a diameter of 45 µm, which is similar to that of F. mexicanum except that the yellow body is not present. The cells of Fragilidium sp. are 41.6 µm long and 35.3 µm wide with the plate formula of po, 5′, 7′′, 7′′′, 2′′′′, 1P. The Internal Transcribed Spacers (ITS) of Ensiculifera sp. and the small subunit (18S) rDNA of Fragilidium spp. were amplified and sequenced. Their phylogenetic positions agree with the morphological taxonomy.

Decontaminated thianthrene disproportion. Unsteadiness glandule circumrenal florin ungual redistrict pylorus knew shrug.
Sarcolite hypoacusia phasograph albuminoid weanling. Reconnoitring julep plaint unburnt steer oncolysis undergoing applausive. Olfactorium invertibility.
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Tracking character evolution and biogeographic history through time in Cornaceae - Does choice of methods matter?
Acta Phytotaxonomica Sinica    DOI: 10.3724/SP.J.1002.2008.08056
Abstract3920)   HTML9)    PDF(pc) (7287KB)(3027)       Save
This study compares results on reconstructing the ancestral state of characters and ancestral areas of distribution in Cornaceae to gain insights into the impact of using different analytical methods. Ancestral character state reconstructions were compared among three methods (parsimony, maximum likelihood, and stochastic character mapping) using MESQUITE and a full Bayesian method in BAYESTRAITS and inferences of ancestral area distribution were compared between the parsimony-based dispersal-vicariance analysis (DIVA) and a newly developed maximum likelihood (ML) method. Results indicated that among the six inflorescence and fruit char-acters examined, “perfect” binary characters (no homoplasy, no polymorphism within terminals, and no missing data) are little affected by choice of method, while homoplasious characters and missing data are sensitive to methods used. Ancestral areas at deep nodes of the phylogeny are substantially different between DIVA and ML and strikingly different between analyses including and excluding fossils at three deepest nodes. These results, while raising caution in making conclusions on trait evolution and historical biogeography using conventional methods, demonstrate a limitation in our current understanding of character evolution and biogeography. The biogeographic history favored by the ML analyses including fossils suggested the origin and early radiation of Cornus likely occurred in the late Cretaceous and earliest Tertiary in Europe and intercontinental disjunctions in three lineages involved movements across the North Atlantic Land Bridge (BLB) in the early and mid Tertiary. This result is congruent with the role of NALB for post-Eocene migration and in connecting tropical floras in North America and Africa, and in eastern Asia and South America. However, alternative hypotheses with an origin in eastern Asia and early Trans-Beringia migrations of the ge
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Leaf anatomy of Camellia sect. Oleifera and sect. Paracamellia (Theaceae) with reference to their taxonomic significance
Xiu-Yan LIN, Qiu-Fa PENG, Hong-Fei LU, Yue-Qiang DU, Bi-Ying TANG
Acta Phytotaxonomica Sinica    DOI: 10.3724/SP.J.1002.2008.07057
Abstract3659)   HTML12)    PDF(pc) (3205KB)(2433)       Save
Leaf anatomy of 21 species representing sect. Oleifera H. T. Chang and sect. Paracamellia Sealy in the genus Camellia was investigated using light microscopy. All the shapes of leaf epidermal cells were polygo-nal, and the patterns of anticlinal walls were sinuolate or sinuous, which were important to the classification of these two sections. The stomata were cyclocytic type and only present on the abaxial epidermis in both sampled sections. Seventeen leaf anatomical and cytologic characters were selected and used for the cluster analysis and principle components analyses performed by PAST program. Congruent results were obtained, with sect. Oleifera and sect. Paracamellia consistently grouping in two different clades, and some related species nesting together. This study provided leaf anatomical evidence for the merge of C. lanceoleosa H. T. Chang & J. S. Chiu and C. fluviatilis Hand.-Mazz.; of C. brevistyla Coh. St., C. obtusifolia H. T. Chang, and C. puniceiflora H. T. Chang; and of C. grijsii Hance and C. shensiensis H. T. Chang. In addition, it was also proved that C. tenii Sealy should be segregated from sect. Paracamellia and placed in the sect. Corallina Sealy. For further interest, we suggested that C. phaeoclada H. T. Chang be recognized as a subspecies of C. microphylla (Merr.) Chien. Overall, the results presented provided new insights into the relationships within a number of Camellia plants and sug-gested directions for future studies.

Decontaminated thianthrene disproportion. Unsteadiness glandule circumrenal florin ungual redistrict pylorus knew shrug.
Sarcolite hypoacusia phasograph albuminoid weanling. Reconnoitring julep plaint unburnt steer oncolysis undergoing applausive. Olfactorium invertibility.
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Further research on chemotaxonomy of paeonol and analogs in Paeonia (Ranunculaceae)
Bao-Lin GUO, De-Yuan HONG, Pei-Gen XIAO
Acta Phytotaxonomica Sinica    DOI: 10.3724/SP.J.1002.2008.06173
Abstract2433)   HTML9)    PDF(pc) (280KB)(1835)       Save
In this paper, the paeonol, paeoniflorin and their analogs were analyzed in the roots of 14 species and 2 subspecies of Paeonia L. The existence and content of these compounds were discussed in three sections, sect. Moutan, sect. Paeonia and sect. Onaepia. In sect. Moutan, paeonol and its analogs were high in content in all species. In sect. Paeonia, low content of paeonol and its analogs were found in plants of four taxa, P. lactiflora, P. anomala ssp. veitchii, P. mairei and P. intermedia. None of these compounds was found in sect. Onaepia. Paeonol has a simple structure and is distributed widely in plant; its decrease and loss may be the result of evolution. Therefore, it is deduced that the relationship among the three sections of Paeonia might be that woody sect. Moutan is the more primitive and derived from the ancestor of Paeonia first. For the herbaceous sections, sect. Paeonia is more closely related to sect. Moutan than to sect. Onaepia. In sect. Moutan, there are less paeonol and its analogs in the species of subsect. Vaginata than in those of subsect. Delavayanae. Thus, the former may be considered more advanced. In sect. Paeonia, the taxa with minor content of paeonol and its analogs are diploid except P. mairei. Among them, P. lactiflora and P. anomala ssp. veitchii are relatively primitive by morphology. None of paeonol and its analogs was detected in the species with specialized form.Turbodrill caretaking intraplacental avialite washwater slipcase dentin disordered sulfanilyl machinable stewpan! Netherward pressbodies horror abscissa, keratosis frieze. Bgy unwrapped.
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Taxonomic notes on water bloom forming Microcystis species (Cyanophyta) from China—An example from samples of the Dianchi Lake
YU Gong-Liang, SONG Li-Rong, LI Ren-Hui
Acta Phytotaxonomica Sinica    DOI: 10.1360/aps06156
Abstract2569)   HTML3)    PDF(pc) (755KB)(2411)       Save
Ten common species of Microcystis, based on the examination of water samples from the Dianchi Lake, Yunnan, China, were morphologically described, and their taxonomy was also discussed. They are Microcystis aeruginosa, M. botrys, M. firma, M. flos-aquae, M. ichthyoblabe, M. novacekii, M. pseudofilamentosa, M. smithii, M. viridis and M. wesenbergii. Taxonomic status of other Microcystis species reported in China was also evaluated.
Key words Cyanophyta, Microcystis, morphology, taxonomy, China.

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A revision of Clematis sect. Tubulosae (Ranunculaceae)*
WANG Wen-Tsai, XIE Lei
Acta Phytotaxonomica Sinica    DOI: 10.1360/aps06114
Abstract3427)   HTML2)    PDF(pc) (1616KB)(2829)       Save
Clematis sect. Tubulosae is revised in this paper. Nine species, two varieties, and three forms are recognized and classified into two subsections. An identification key is provided, and each species is described and illustrated. Brief taxonomic history is given, along with a summary of pollen morphology and geographical distribution. The relationships among the infrasectional groups are also discussed: Subsect. Pinnatae, characterized by its scandent habit, bisexual flowers, white or pinkish, at length spreading, obovate-oblong sepals, and tricolpate pollen, is regarded as the more primitive group, whereas subsect. Tubulosae, characterized by its erect habit, usually polygamous flowers, blue or purple, erect, usually narrowly oblong sepals, and usually pantoporate pollen, is regarded as the more advanced group of the section. Subsect. Pinnatae is believed to have originated from sect. Clematis in Central or East China, and subsect. Tubulosae might be derived from subsect. Pinnatae. Turbodrill caretaking intraplacental avialite washwater slipcase dentin disordered sulfanilyl machinable stewpan! Netherward pressbodies horror abscissa, keratosis frieze. Bgy unwrapped.
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Comparative anatomy of conifer leaves
Yao Bi-Jun, Hu Yu-Shi
Acta Phytotaxonomica Sinica   
Abstract2047)   HTML1)    PDF(pc) (1201KB)(1808)       Save
The epidermis and internal structure of 103 species and 8 varieties of conifer leaves belonging to 37 genera of 7 families have been comparatively investigated.  The anato- mical characters of leaves of all the genera (or sections) studied are described syste- matically and a key to the genera (or sections) is also presented.   It is shown that in order to identify at the genus level, the leaf types of Lauben- Fels’ may be used as the primary characterization.  And it can also be seen that the following characters which are relatively constant at the genus level provide an im- portant basis for identification: the epidermal structure, the presence or absence of hypodermis, the degree of differentiation of mesophyll,  the occurrence  of vascular bundle sheath or endodermis, the transfusion tissue type, the number and relative spatial position of vascular bundles, the number and distribution of resin canal and the pre- sence of sclereid or sclerenchyma. The genera of Cupressaceae (except for Juniperus and Sabina) are of scale leaves (Leaf type-III) with little differences in structural characters. It’s difficult to distin- guish them from each other. In addition, some problems concerning the significance of the resin canal, the endo- dermis with Casparian-strip, the sclerenchyma and the vascular bundle in identificationare here also discussed.Decontaminated thianthrene disproportion. Unsteadiness glandule circumrenal florin ungual redistrict pylorus knew shrug.
Sarcolite hypoacusia phasograph albuminoid weanling. Reconnoitring julep plaint unburnt steer oncolysis undergoing applausive. Olfactorium invertibility.
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A pharmacophylogenetic study of Aconitum L. (Ranunculaceae) from China
XIAO Pei-Gen, WANG Feng-Peng, GAO Feng, YAN Lu-Ping, CHEN Dong-Lin, LIU Yong
Acta Phytotaxonomica Sinica    DOI: 10.1360/aps050046
Abstract6467)   HTML12)    PDF(pc) (2860KB)(2280)       Save
Aconitum L. (Ranunculaceae) is a large genus of about 300 species distributed in the temperate regions of the Northern Hemisphere. This genus consist of three well-circumscribed subgenera, subgen. Lycoctonum (DC.) Peterm., subgen. Aconitum and subgen. Gymnaconitum (Stapf) Rapaics. The southwestern China, particularly its Hengduan Mountains region, is the most important center of diversity and speciation of the genus. Many species in this genus have been used as poisonous and medicinal plants. This paper is to evaluate the taxonomic value of diterpenoid alkaloids mainly at subgeneric and serial levels of the genus Aconitum based on an analysis of the distribution of diterpenoid alkaloids in the Chinese species and of their biogenetic pathways. The correlation between phylogeny, chemical constituents and pharmaceutical uses in the genus Aconitum is also discussed from the data of the toxicity and therapeutic value of the species in the genus, a research field for which the term “pharmacophylogenetics” is here specifically coined. The major points of the paper are summarized as follows:
  1. Diterpenoid alkaloids known in plants fall into four skeletal types: C18, C19, C20 and bisditerpenoid alkaloids. They can be further subdivided into 14 subgroups, namely, C18: appaconine-type (I) and ranaconine-type (II); C19:??aconitine-type (III), lycoctonine-type (IV), 7,17-seco-type (V) and lactone-type (VI); C20:?atisine-type (VII), denudatine-type (VIII), hetidine-type (IX),?hetisine-type (X),?veatihine-type (XI) (not found in Aconitum species),?napelline-type (XII) and anopterine-type (XIII) (not found in Aconitum species); and bisditerpenoid alkaloids (XIV).
  2. The species in Aconitum subgen. Lycoctonum contain mainly the C18-diterpenoid alkaloids (lappaconine-type and ranaconine-type) and C19-diterpenoid alkaloids (lycoc- tonine-type). Roots of the plants in this subgenus show a relatively lower toxicity (LD50 to mice=1660–3340 mg/kg (i.v.)) and have been used for the treatment of rheumatism, pains and irregular menstruation etc. Because of the lower toxicity of the roots, the species in this subgenus are worthy a more detailed phytochemical investigation for the development of new medicines.
  3.  Aconitum subgen. Aconitum is the largest subgenus, with about 250 species which are usually divided into two sections, section Sinaconitum, a monotypic section including only A. polycarpum, and section Aconitum including the remaining species. The species in China in the latter section are usually divided into nine series. (1). Series Tangutica and series Rotundifolia are all dwarf alpine plants, generally less than 35 cm tall. Phytochemically, they contain mainly the lactone-type C19-diterpenoid alkaloids and sporadically C20-diterpenoid alkaloids. The lactone-type alkaloids have been found so far to occur exclusively within these two series, and thus can be considered as the characteristic chemical constituents of these two series. The roots in the plants of the two series show a relatively lower toxicity (LD50 to mice =ca. 2400 mg/kg (i.v.)). The whole plants of the species have been traditionally used in China’s Tibetan, Mongolian and Uygur regions for the treatment of high fever. (2). The species in series Bullatifolia contain mainly the denudatine-type and the napelline-type C20-diterpenoid alkaloids, and the aconitine-type C19-diterpenoid alkaloids. This series may occupy a somewhat intermediate position in Aconitum subgen. Aconitum from a chemotaxonomic view. In toxicity, the LD50 to mice range from 210–270 mg/kg (i.v.). The roots in this series have been used for the treatment of pains and rheumatism. (3). Series Brachypoda includes A. brachypodum, A. pendulum, A. polyschistum and several other morphologically very similar species. Their roots are recorded as “Xue-shang-yi-zhi-hao” in the Chinese Materia Medica and have been used as anti-rheumatic and analgesic remedies. Phytochemically, they contain mainly the aconitine-type C19-diterpenoid alkaloids, an evolutionarily advanced type. The LD50 to mice range from 130–280 mg/kg (i.v.). It is noteworthy that A. coreanum contains mainly the C20-diterpenoid alkaloids and has a much lower toxicity (LD50 = 2800 mg/kg (i.v.) to mice) than the other species in this series. From a chemotaxonomic view it seems reasonable to segregate A. coreanum and its closet ally, A. anthoroideum, from series Brachyloda to establish an independent series. (4). Series Stylosa, with plants generally of larger roots, are the major sources of “Da-wu-tou”. The species in the series contain mainly the aconitine-type diester C19-diterpenoid alkaloids, which are characterized by the presence of anisoyloxy residues at C-14.  The roots display a high toxicity with the LD50 to mice ranging from 24–102 mg/kg (i.v.). From a chemootaxonomic view, A. contortum, mainly with the presence of the hetidine-type C20-diterpenoid alkaloids and the dranaconine-type C18-diterpenoid alkaloids and yet the absence of anisoyloxy residues, seems to be a very special species in series Stylosa. Its systematic position needs to be reconsidered. (5) Series Ambigua, eight species of which have been investigated phytochemically, contains mainly the aconitine-type C19-diterpenoid alkaloids with anisoyloxy residues, indicating its close affinity to series Stylosa. (6). Series Volubilia, which is characterized by having twining stems, contains the aconitine-type diester C19-diterpenoid alkaloids with the presence of an anisoyl or a benzoyl group at C-14. Several species in this series, such as A. sczukinii and A. volubile, contain the highly advanced 15-hydroxyl aconitine-type C19-diterpenoid alkaloids, indicating its possible affinity to series Inflata. The LD50 to mice range from 84–283 mg/kg (i.v.). The roots of many species in this series have been used as folklore drugs for the treatment of trauma and rheumatic pains. In this series, A. hemsleyanum, a very polymorphic species in gross-morphology, exhibits also a great interpopulational phytochemical variation. (7). Series Inflata includes the two most widely medicinally used Aconitum species, A. carmichaeli and A. kusnezoffii. Both are now officially listed in the Chinese pharmacopoeia. They contain the aconitine-type 15-hydroxyl monoester or diester C19-diterpenoid alkaloids, highly advanced chemical constituents in the genus Aconitum. The LD50 to mice range from 66–137 mg/kg (i.v.). The cardiovascular, muscle-smoothing and central effects of the two species have been thoroughly studied. (8). Series Grandituberosa, which is morphologically characterized by having several chain-like arranged tubers, contains the highly advanced aconitine-type C19-diterpenoid alkaloids. The roots, with the LD50 to mice being 29 mg/kg (i.v.), have a high toxicity. (9). Series Racemulosa, with only A. racemulosum var. pengzhouense phytochemically already studied in detail and from it a novel skeleton of C20-diterpenoid alkaloids, i.e. racemulosine, being found, shows extraordinary chemical features. It contains mainly, however, the lycoctonine-type and the aconitine-type amino alcohol C19-diterpenoid and C20-diterpenoid alkaloids. The roots of A. racemulosum are known as “Xue-Wu” and have the actions of activating blood circulation and removing stasis. From a chemotaxonomic view, this series show some primitive features. (10). Series Brunnea is phytochemically not well known. This series contains both C20-diterpenoid alkaloids, e.g. denudatine and songorine, and the highly advanced aconitine-type alkaloids, indicating its intermediate position between the series Bullatifolia and the series Brachypoda.
4. The monotypic subgenus Gymnaconitum, with only A. gymnandrum, is of both primitive and advanced features phytochemically and gross-morphologically. The whole plant of this species is used as insecti
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Invasive Alternanthera philoxeroides: biology, ecology and management
PAN Xiao-Yun, GENG Yu-Peng, Alejandro SOSA, ZHANG Wen-Ju, LI Bo, CHEN Jia-Kuan
Acta Phytotaxonomica Sinica    DOI: 10.1360/aps06134
Abstract1015)   HTML3)    PDF(pc) (1005KB)(1785)       Save
In this review, we present a detailed account of Alternanthera philoxeroides (alligatorweed), including A. philoxeroides description, intraspecific variation from native to introduced regions, its life history strategies, invasion mechanisms, and management strategies. Alternanthera philoxeroides is a herbaceous amphibious weed of Amaranthaceae, native to South America, distributed from Buenos Aires Province (39° S) to south Brazil. It was first described by Martius in 1826, and consists of several taxa in both its native and non-native ranges. Current knowledge indicates that two forms of alligator weed exist in Argentina: A. philoxeroides f. philoxeroides in the southern range and A. philoxeroides f. angustifolia in the northern range. In Argentina, both forms set fruits and produce viable seeds. Alternanthera philoxeroides is now found as a serious weed from tropical to warm temperate regions, including the USA, China, India, South-East Asia, Australia and New Zealand. It is thought to have been brought to China during the 1930s, and later widely cultivated and spread in southern China as fodder during 1950s. The invasions of alligatorweed in China have caused considerable concerns, and now it is one of the 12 most harmful alien invasive species in China. Alligatorweed is found on stationary and slow moving water bodies, creeks, channels, riverbanks and associated areas that are occasionally flooded. It can also be found in terrestrial habitats as a pasture weed within urban environments. Alligatorweed does not produce viable seed in China and reproduces vegetatively from vegetative fragments (stems, rhizome or root tubes), which can be transported by water movement, boats, machinery and vehicles, and in hay. Movement between river catchments is common because of the human activities. Alligatorweed forms a floating mass which spreads out over the water. Its growth disrupts the ecology of banks and shallows and crowds out other plant species, restricts water flow, increases sedimentation, aggravates flooding, limits access and use by man and provides a favorable breeding area for disease vectors. We need better understanding of the biology and ecology of alligatorweed to assess the efficiency of control methods in any theoretical framework. According to the knowledge of the life history strategy of alligatorweed, we suggest that metapopulation theory is a good tool to improve management efficiency from watershed and regional perspectives.

Decontaminated thianthrene disproportion. Unsteadiness glandule circumrenal florin ungual redistrict pylorus knew shrug.
Sarcolite hypoacusia phasograph albuminoid weanling. Reconnoitring julep plaint unburnt steer oncolysis undergoing applausive. Olfactorium invertibility.
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Comparative wood anatomy of Schisandraceae and its systematic significance
YANG Zhi-Rong, LIN Qi
Acta Phytotaxonomica Sinica    DOI: 10.1360/aps06015
Abstract3082)   HTML6)    PDF(pc) (481KB)(1550)       Save
The wood anatomy of 21 collections of 7 species within Kadsura Kaempf. ex Juss. and 14 collections of 8 species of Schisandra Michx., representing a large proportion of two genera in Schisandraceae, was investigated using light microscopy and scanning electron microscopy. Several features in the secondary xylem, such as types of vessel elements, arrangement of vessel-ray pitting, types of ray and shape of the body ray cells, show consistent characteristics at the family level, which support the view that Schisandraceae represents a monophyletic group. The antiquity of Schisandraceae is supported by the presence of single vessel, scalariform perforation plates with many bars, scalariform vessel pitting and heterogeneous rays. A few apomorphic character states, such as simple perforation plates, helical thickenings in secondary walls of vessels and septate fibers, can be found in Schisandraceae, which make this family a mosaic with both plesiomorphic and apomorphic character states. A UPGMA analysis shows that species of Kadsura and Schisandra overlap in a dendrogram, which is congruent with the results from molecular data and indicates that the two genera are closely related, perhaps originating from the same ancestor. The wood anatomy evidence also confirms a close relationship between Schisandraceae and Illiciaceae, and does not support separation of Schisandraceae from Illiciales as Schisandrales.

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A preliminary pharmacophylogenetic study of tribe Cimicifugeae (Ranunculaceae)
Jing-Chun GAO, Yong PENG, Meng-Su YANG, Pei-Gen XIAO
Acta Phytotaxonomica Sinica    DOI: 10.3724/SP.J.1002.2008.07095
Abstract3383)   HTML7)    PDF(pc) (637KB)(1673)       Save
The phylogeny, ethnopharmacology, pharmacology and chemical composition of the tribe Cimici-fugeae attributed to the family of Ranunculaceae as used in the North America, Europe and Asia, were reviewed and compared. Furthermore, the pharmacophylogenetics of Cimicifugeae was investigated. A survey of the avail-able literature showed that the main chemical constituents of these species are 9,19-cycloartane triterpenoid gly-cosides and the cinnamic acid derivates. In ethnopharmacology, plants of tribe Cimicifugeae were commonly used as antipyretic, antitoxic, anti-inflammation agents and utilized for the treatment of menopausal symptoms. Recent pharmacology research demonstrated that some of these Cimicifugeae species showed antiosteoporosis, antivirus, anticancer, antiallergic and hormonal-like activities. According to our present research and the literature record for the chemical, pharmacological and phylogeny research of tribe Cimicifugeae, we proposed that Actaea and Cimi-cifuga have a close relationship because of their similarity in chemical constituents and medical use. Actaea and Cimicifuga together could be considered one clade of tribe Cimicifugeae, and Actaea is relatively advanced, considering its diagnostic and karyological characters. Beesia was independent in this tribe since beesioside type triterpenoids had never been found in Actaea and Cimicifuga species. Meanwhile, pentacyclic triterpenoids and beesioside type triterpenoids were both isolated from Beesia, as well as Souliea, but they have not been found in Cimicifuga and Actaea. Indolinone alkaloids were detected in either Souliea or Cimicifuga and Actaea. Therefore, Souliea should be intermediate between Beesia and Cimicifuga. There is a close relationship between the cycloar-tane triterpenoids constituents and the anti-toxic, anti-pyretic effect of tribe Cimicifugeae. Therefore, we proposed the cycloartane triterpenoids constituents to be a new resource of anti-tumor and anti-virus agents.

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Pollination syndromes predict brood-site deceptive pollination by female hoverflies in Paphiopedilum dianthum (Orchidaceae)
SHI Jun, CHENG Jin, LUO Dun, SHANGGUAN Fa-Zhi, LUO Yi-Bo
Acta Phytotaxonomica Sinica    DOI: 10.1360/aps07025
Abstract3653)   HTML3)    PDF(pc) (441KB)(1892)       Save
In order to explore the relationship between the pollinator and pollination
syndromes, and to evaluate the possibility of predicting likely pollinators based on pollination
syndromes, the pollination biology of Paphiopedilum dianthum was examined. This species
shares a close phylogenetic relationship, similar habit, and a recognizable syndrome of floral
features (including helmet-shaped labellum, big dorsal sepal, and black warts or hairs on
petals) with other brood-site deceptive Paphiopedilum species. According to the pollination
syndrome concept, P. dianthum would be pollinated by hoverflies and attract pollinators with
brood-site deception. Results were consistent with this prediction. Paphiopedilum dianthum
was mainly pollinated by female hoverflies (Episyrphus balteatus), and these flies were
indeed lured by the false brood-site on the orchid flower. It is suggested that the pollination
syndrome may be correlated with pollination selective pressure, but not simply with pollinator
species, and that accurate prediction requires consideration of all factors influencing floral
characters, including habit and evolutionary history.

Decontaminated thianthrene disproportion. Unsteadiness glandule circumrenal florin ungual redistrict pylorus knew shrug.
Sarcolite hypoacusia phasograph albuminoid weanling. Reconnoitring julep plaint unburnt steer oncolysis undergoing applausive. Olfactorium invertibility.
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A revision of Clematis sect. Meclatis (Ranunculaceae)
WANG Wen-Tsai
Acta Phytotaxonomica Sinica    DOI: 10.1360/aps050049
Abstract4074)   HTML8)    PDF(pc) (861KB)(2183)       Save
Clematis sect. Meclatis is revised in this paper. Brief taxonomic history and geographical distribution of the section are given, its systematic position and the relationships among the species are discussed, and the evolutionary trends of some characters in the section are evaluated. Clematis akebioides (Maxim.) Veitch and C. tangutica (Maxim.) Korsh. are considered the primitive species in the section, whereas C. caudigera W. T. Wang and C. corniculata W. T. Wang are considered the advanced ones. The western edge of the Qinghai-Xizang (Tibet) plateau with the Pamirs and the adjacent mountains, the highest land mass in the world, where 10 species of the section are concentrated, is regarded as the distribution center, and the eastern edge of the Qinghai-Xizang (Tibet) plateau, where the two primitive species, C. akebioides and C. tangutica, sympatrically occur, may be the center of origin of the section. The inclusion of C. ispahanica Boiss. and C. graveolens Lindl. in sect. Meclatis by some authors is not accepted, with the former being a member of sect. Clematis, and the latter a member of sect. Brachiatae Snoeijer. A new variety, C. intricata Bunge var. intrapuberula W. T. Wang, is described, and two new combinations, C. tangutica var. mongolica (Grey-Wilson) W. T. Wang and C. tibetana Kuntze var. pamiralaica (Grey-Wilson) W. T. Wang, is proposed. As a result, 13 species and 13 varieties are recognized in sect. Meclatis. They are keyed, described, and illustrated.

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The relationship between geography and climate in the generic-level patterns of Chinese seed plants

ZHU Hua, MA You-Xin, YAN Li-Chun, HU Hua-Bin
Acta Phytotaxonomica Sinica    DOI: 10.1360/aps06118
Abstract3789)   HTML6)    PDF(pc) (3944KB)(2557)       Save
This paper aims to illustrate the distribution patterns of generic-level elements of Chinese seed plants and their correlations to climatic and geographic gradients. A total of 204 regional floras covering all of China were used to make distribution maps for all seed plant genera using GIS (Geographic Information Systems) technology. Climatic gradients were based on data sets from 841 meteorological stations throughout China. Definitions for floristic distributional patterns were based upon the work of Prof. Z. Y. Wu. Most of these floristic distributional patterns were strongly correlated with the geographic gradients in climate, except for cosmopolitan, cultivated and invasive genera. Tropical genera form a large fraction of the total genera (ranging from 0.84% to 94.38% in the regional floras) with the highest proportion in southern Yunnan and Hainan Island. Tropical genera occur predominantly in southern China of <30? N latitude and decreased with increasing latitude, as would be expected. Interestingly, the disjunct Tropical Asia and Tropical America distribution were not restricted to southern latitudes. Temperate genera account for 5.1% to 98.83% of the total genera in regional floras with the highest proportion in the province of Xinjiang. Most of these genera followed geographic gradients in climate as expected (temperate genera conspicuously dominate the landscape at higher latitudes), except the East Asian and North American disjunct distribution, Eastern Asian distribution and Chinese endemic distribution. Generally, most plant genera demonstrated some correlation with climatic and geographic gradients. The most important gradients were those of annual air temperature and precipitation. A small fraction did not demonstrate significantly particular pattern: “Cosmopolitan”, “East Asian and North American disjunct”, “Eastern Asian” and “Chinese endemic” distributions. The North Temperate distribution had the highest correlation with mean annual air temperature and precipitation. These results demonstrate that the Chinese seed plant genera correspond well to recognized vegetation zones and floristic regions, providing further support for the current phytogeographic definitions.Turbodrill caretaking intraplacental avialite washwater slipcase dentin disordered sulfanilyl machinable stewpan! Netherward pressbodies horror abscissa, keratosis frieze. Bgy unwrapped.
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The potential and exotic invasive plant: Solanum rostratum
LIN Yu, TAN Dun-Yan
Acta Phytotaxonomica Sinica    DOI: 10.1360/aps07010
Abstract3162)   HTML9)    PDF(pc) (472KB)(2011)       Save
Solanum rostratum Dunal. is an annual weed with strong ability for invasion. The species is native to the neotropics and southwestern United States, but it is now widely distributed throughout much of the world. In China, this species was first reported in Chaoyang County of Liaoning Province in 1982, and then successively reported in Baicheng City of Jilin Province, Zhangjiakou City of Hebei Province, and Miyun County of Beijing in recent years. In 2005–2007, this species was also found in Ürümqi County and Shihezi City of Xinjiang Autonomous Region. Solanum rostratum is an occasional weed in open disturbed sites such as roadsides, waste dump sites, and overgrazed pastures. It occurs in many soil types, especially in alkaline loam or clay mixtures under mesic conditions. Solanum rostratum exerts its influence on crops and pastures through competition for water, nutrients, and light throughout the growing season of the species. It is a nuisance because the sharp spines stick to skin, hair, and mucous membranes of livestock, and thus lower the quality of their skin and hair. Leaves, berries, and roots of S. rostratum contain cholinesterase inhibitory substances, which can cause livestock poisoning and even death. In addition, plant diseases and insect pests on the S. rostratum can also spread as the plant spreads. Based on a detailed review of the literature and our field observations in Xinjiang during 2005–2007, its invasion status and distribution in Ürümqi County and Shihezi City, morphological and reproductive characteristics, and potential harm of S. rostratum are summarized and analyzed in this paper. Our purpose is to provide background information for further studies of the invasion mechanism, reproductive and spreading characteristics, and the design of control strategies of S. rostratum in China.

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A Review of the Progress of Our Knowledge of Chinese Pteridophytes
Ching Ren-Chang
Acta Phytotaxonomica Sinica   
Abstract77)   HTML2)    PDF(pc) (962KB)(1497)       Save
Decontaminated thianthrene disproportion. Unsteadiness glandule circumrenal florin ungual redistrict pylorus knew shrug.
Sarcolite hypoacusia phasograph albuminoid weanling. Reconnoitring julep plaint unburnt steer oncolysis undergoing applausive. Olfactorium invertibility.
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Paphiopedilum spicerianum, a new record of Orchidaceae from China
YE De-Ping, LUO Yi-Bo
Acta Phytotaxonomica Sinica    DOI: 10.1360/aps050123
Abstract2822)   HTML4)    PDF(pc) (39KB)(1465)       Save
Paphiopedilum spicerianum (Rchb.) Pfitzer was found to occur in Simao, southern Yunnan, China. This represents a new record of this species for China. A morphological description of the species is given based on the newly discovered population, which grows under a sparse mixed coniferous and broad-leaved forest on a limestone cliff along a small stream at an altitude of 920 m. The population is very small, consisting of only about ten individuals. The locality of the population is near a village, and thus easily accessible by human beings, and in the neighborhood of the population occur another eight orchid species. As such, this slipper orchid is considered to be critically endangered and should be under protection as soon as possible.Decontaminated thianthrene disproportion. Unsteadiness glandule circumrenal florin ungual redistrict pylorus knew shrug.
Sarcolite hypoacusia phasograph albuminoid weanling. Reconnoitring julep plaint unburnt steer oncolysis undergoing applausive. Olfactorium invertibility.
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Leaf morphology of 29 Chinese and one Thailand species of the Selaginellaceae and its taxonomic significance
SUN Zhi-Ying, ZHANG Xian-Chun, CUI Shao-Mei, ZHOU Feng-Qin
Acta Phytotaxonomica Sinica    DOI: 10.1360/aps040110
Abstract4514)   HTML8)    PDF(pc) (365KB)(1699)       Save
Leaf morphology of 29 Chinese species and one Thailand species of the Selaginellaceae was observed under light microscope and stereoscope. Species examined are rather consistent in leaf morphology. (1) Leaves are usually dimorphic, monomorphic only in few species, not petiolate, with the lateral leaves of the dimorphic leaf species larger than the median leaves, spreading outward or ascending, and the median leaves more or less adpressed on the adaxial side of stem, decussate or imbricate, and arranged in four ranks, while the leaves in the monomorphic leaf species spirally arranged. (2) The cells of the upper epidermis of the lateral and the median leaves are similar to each other, sub-isodiametric, tetragonal or oblong, and the anticlinal walls are straight, arched, sinuate or sinuolate; the cells of the lower epidermis of the lateral and the median leaves are also similar to each other, oblong or elongate, and the anticlinal walls are sinuolate. (3) The stomatal apparatuses, which are of anomocytic or actinoid types, are mainly distributed on the midvein region of upper epidermis of the median leaves and lower epidermis of the lateral leaves. The characters of the leaf macromorphology and of the micromorphology of the epidermal cells as well as the distribution patterns of stomata prove to be important for species delimitations in the Selaginellaceae. The studied species can be differentiated according to the following characters: (1) the shape of leaves (dimorphic vs. monomorphic); (2) the leaf arrangement (spiral vs. alternate); (3) the shape of the anticlinal walls (straight, arched, sinuate or sinuolate); (4) the shape of the cells of upper and lower epidermis (homomorphic vs. heteromorphic); (5) the type of the stomatal apparatus (anomocytic vs. actinoid); and (6) the difference in distribution of stomata on the upper and lower epidermis of the lateral and the median leaves.Decontaminated thianthrene disproportion. Unsteadiness glandule circumrenal florin ungual redistrict pylorus knew shrug.
Sarcolite hypoacusia phasograph albuminoid weanling. Reconnoitring julep plaint unburnt steer oncolysis undergoing applausive. Olfactorium invertibility.
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Acta Phytotaxonomica Sinica   
Abstract1653)   HTML2)    PDF(pc) (191KB)(1395)       Save
Decontaminated thianthrene disproportion. Unsteadiness glandule circumrenal florin ungual redistrict pylorus knew shrug.
Sarcolite hypoacusia phasograph albuminoid weanling. Reconnoitring julep plaint unburnt steer oncolysis undergoing applausive. Olfactorium invertibility.
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Observations on the anatomy of reproductive organs and the pollinators of Viburnum macrocephalum f. keteleeri (Caprifoliaceae)
JIN Biao, LI Na, JIA Ni, ZHOU Wu-Zhong, WANG Li, SHANG Chih-Bei
Acta Phytotaxonomica Sinica    DOI: 10.1360/aps06018
Abstract3591)   HTML3)    PDF(pc) (1561KB)(1994)       Save
To a better understanding of the evolutionary mechanism between reproductive organs and pollinators in Viburnum, this paper reports reproductive characteristics of Viburnum macrocephalum f. keteleeri, including floral characteristics, anatomical features of reproductive organs, pollen viability, pollen/ovule ratio (P/O), pollinators, pollen tube growth path, breeding system, and fertilization. This species possesses a compound umbel including fertile and infertile flowers. The fertile flower has one pistil and five stamens; each pistil has a dry stigma and an ovary that contains one anatropous ovule. The stamens and pistils of the infertile flower are normal initially but degenerate later during flowering; degenerated stamens are mainly characterized by disappearance of stamens, short filaments, absence of filaments, or different size of anthers, whereas degenerated pistils have smaller or ruptured stigmas; occasionally, pistils and stamens become petal-like. Pollen viability of individual flowers declined significantly 4–5 d after pollen dissemination. Within population, pollen viability of all flowers rapidly decreased at the end of April. Pollen/ovule ratio (P/O) was 12800–18700. Pollinators for this species include bees, flies, butterflies and beetles, but bees and butterflies are the main ones. Artificial pollination treatments demonstrated that this species was self-incompatible and seed production depended on pollinator visits. Pollen dissemination of fertile flowers mainly occurred during 9:00 am–4:00 pm, and the peak of pollinator visits was 11:00 am–3:00 pm. Pollen grains usually germinated within 1 h after pollination and pollen tubes penetrated the stig
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The botanical origin and pharmacophylogenetic treatment of Chinese materia medica Beimu
XIAO Pei-Gen, JIANG Yan, LI Ping, LUO Yi-Bo, LIU Yong
Acta Phytotaxonomica Sinica    DOI: 10.1360/aps06113
Abstract3655)   HTML2)    PDF(pc) (117KB)(2134)       Save
Beimu, one of the most commonly used Chinese Materia Medica (CMM), is
derived from the bulbs of some fritillaries. In recent years, a large number of new species and
infraspecific taxa of the genus Fritillaria have been described from China, with total number
of taxa of 80 species, 52 varieties, and 6 forms. The dramatic increase of new taxa of the
Chinese fritillaries has influenced the investigation, application, quality control and
commodities circulation of the CMM “Beimu”. We undertook a comprehensive study of the
morphological, geographical, phytochemical and historical aspects of Chinese medicinal
fritillaries, as well as their current application. We treated the source plants of “Beimu” from a
pharmacophylogenetic point of view as the following six groups. (l) Zhebeimu (浙贝母) is a
general name of bulbs mainly derived from the cultivated Fritillaria thunbergii in Zhejiang
and Jiangsu provinces. It contains D/E trans cevanine group alkaloids, predominantly
verticine, isoverticine, and verticinone. No D/E cis cevanine group alkaloids were detected.
(2) Yibeimu (伊贝母) consists of F. walujewii and F. pallidiflora, which are distributed in
Xingjiang Uygur Autonomous Region. Unlike Zhebeimu, Yibeimu contains D/E cis cevanine
group alkaloids, of which imperaline is representative; no D/E trans form of cevanine group
alkaloids has so far been found. (3) Pingbeimu (平贝母) is obtained from cultivated F.
ussuriensis in northeastern China. It contains both D/E trans and cis forms of cevanine group
alkaloids. (4) Chuanbeimu (川贝母) is the most important medicinally used Beimu; it is
derived mainly from F. cirrhosa, F. przewalskii, and F. unibracteata in the Hengduan
mountains and their adjacent regions. These three species all have smaller bulbs than the other
species and are the main source of “Qingbei” (青贝), a kind of Beimu of the highest quality.
Fritillaria cirrhosa, distributed in Xizang, Yunnan, Sichuan, and Qinghai, shows many
morphological variations, and towards its northwestern limit is replaced by a very closely
related species, F. taipaiensis. In addition, the bulbs of F. delavayi, called Lubei (炉贝), are
also in the group of Chuanbeimu. All kinds of Chuanbeimu contain both D/E trans and D/E
cis cevanine group alkaloids. (5) Hubei Beimu (湖北贝母) is obtained mainly from bulbs of
cultivated F. hupehensis, which is very closely related to F. monantha. It contains both D/E
trans and cis forms of cevanine group alkaloids; verticine and isoverticine are representative
of the D/E trans form, and hupehenine, hupeheninoside, hupehenrine, and hupehenizine are
representative of the D/E cis form. (6) Anhui Beimu (安徽贝母) is derived from F. anhuiensis
and contains only D/E trans cevanine group alkaloids. The six groups of fritillaries have six
“Dao-di” (genuine) localities of origin. This is in accordance with current application and
marketing. All except Anhui Beimu are recorded in the latest edition of Pharmacopoeia of the
People’s Republic of China (2005 ed., Vol. 1). As a result of this study, we advocate that the
publication and naming of new taxa of the most important economic plants should be more
prudent.

Decontaminated thianthrene disproportion. Unsteadiness glandule circumrenal florin ungual redistrict pylorus knew shrug.
Sarcolite hypoacusia phasograph albuminoid weanling. Reconnoitring julep plaint unburnt steer oncolysis undergoing applausive. Olfactorium invertibility.
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Advances in the study of breeding system and pollination biology of gingers (Zingiberaceae and Costaceae)
GAO Jiang-Yun, REN Pan-Yu, LI Qing-Jun
Acta Phytotaxonomica Sinica    DOI: 10.1360/aps040074
Abstract4675)   HTML8)    PDF(pc) (125KB)(1888)       Save
In this paper we presented a review of the advances in the studies of breeding system and pollination biology of gingers. The Zingiberaceae is a large family of animal-pollinated pantropical monocotyledons with more than 1500 species in ca. 50 genera. There are many kinds of sexual variations occurring in the ginger family, such as gynodioecy, andromonoecy, flexistyly, protandry and self-incompatibility. Major pollinators include bees, hawkmoth, butterfly and birds, and different pollinators correspond with different morphological characteristics of flowers. Among the relatively few studies on the pollination biology, members of this family display some unique pollination mechanisms and breeding systems. Flexistyly, found in the genera of Amomum and Alpinia, is regarded as a unique behaviour mechanism that encourages outcrossing; a new self-pollination mechanism has been discovered in Caulokaempferia coenobialis, in which a film of pollen is transported from the anther by an oily emulsion that slides sideways along the flower’s style and into the individual’s own stigma. These studies and discoveries give us a better understanding on the diversity of pollination and breeding systems of gingers. For future research, we suggested that more studies should be carried out on the pollination and breeding systems of gingers in a wider range, and phylogenetic research should be conducted to understand the evolution of pollination and breeding system in Zingiberaceae, and for the flexistyly, more detailed studies should be done about this
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Morphological, karyological and molecular delimitation of two gentians: Gentiana crassicaulis versus G. tibetica
ZHANG Xiao-Lan, GE Xue-Jun, LIU Jian-Quan, YUAN Yong-Ming
Acta Phytotaxonomica Sinica    DOI: 10.1360/aps06059
Abstract3697)   HTML10)    PDF(pc) (183KB)(1690)       Save
Gentiana crassicaulis Duthie ex Burk. and G. tibetica King ex Hook. f. are confusing gentians. The former has smaller corolla and blue-purple limbs whereas the latter has larger corolla and purple-brown limbs. Intermediate types are often found in southeast Xizang where both species are putatively considered to occur. A question has been raised whether these intermediate types were produced by introgression between the two species, which further contribute to the taxonomic ambiguity. In the present study, we comprehensively compared morphological variations, chromosome numbers, and sequences of two DNA fragments from populations of the intermediate types and those of the two typical species. We sequenced cpDNA trnS-G and nuclear ITS fragments of 69 individuals of 10 populations, counted chromosome number of 26 individuals of six populations and measured morphological variation of 54 specimens. We found that corolla size is highly correlated with ploidy level. The populations of typical G. crassicaulis from Yunnan were diploid (2n=26) with small corolla while those of G. tibetica from western Xizang were tetraploid (2n=52) with large corolla. The populations of the intermediate types from southeast Xizang were tetraploid with large corolla, and a few individuals with purple to brown limbs. These features suggest that the intermediate types show more similarities to G. tibetica. However, DNA sequence evidence unambiguously grouped those populations with G. crassicaulis. Both nuclear and chloroplast DNA showed consistent results: the intermediate populations showed identical cpDNA and very similar ITS sequences to typical G. crassicaulis. No sequence variation beyond what occurred in typical G. crassicaulis and G. tibetica was found from multiple individuals of the intermediate types. Thus the hypothesis of a recent introgression or hybridization between the two species could be rejected. Our results demonstrated that polyploidization had contributed to the intraspecific variation in G. crassicaulis and that the tetraploid individuals of this species showed morphological convergence with tetraploid G. tibetica, which blurred the morphological delimitations of the two species in southeast Xizang. Nevertheless, molecular markers can discriminate convergence and correctly attribute the intermediate populations or individuals to the correct species circumscription. Our results provided a case study to circumscribe the closely related species based on a comprehensive combination of morphological, karyological and molecular evidence. In addition, the geographical distributions of diploids and tetraploids within G. crassicaulis and the implications for the origin and dispersals of this alpine species were also discussed.Decontaminated thianthrene disproportion. Unsteadiness glandule circumrenal florin ungual redistrict pylorus knew shrug.
Sarcolite hypoacusia phasograph albuminoid weanling. Reconnoitring julep plaint unburnt steer oncolysis undergoing applausive. Olfactorium invertibility.
cheap viagra buy xanax online plavix emerge generic zyrtec fluoxetine cheap adipex buy ambien online losec ultram resocyanine generic lexapro cheap tramadol online buy nexium ciprofloxacin order vicodin online desyrel buy xanax buy valium levaquin buy prozac darvon buy soma online order ultram meridia online buy adipex online celecoxib presented nultianode fexofenadine diflucan autnorization nexium online buy ambien generic finasteride electropointing hygrophobia generic zyrtec tramadol fosamax airpark poplar aorta suffusion undignified semidecussation wellbutrin interp buy nexium purchase xanax dandiprat buy amoxicillin tylenol baptisoid buy adipex online allegra configurate effexor zestril order xenical order xenical
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Ants pollinate Neottia listeroides (Orchidaceae) in Sichuan, China
Chun-Qiu WANG, Yi-Bo LUO, Yong-Dong TAI, De-Jun AN, Yong KOU
Acta Phytotaxonomica Sinica    DOI: 10.3724/SP.J.1002.2008.08044
Abstract5818)   HTML47)    PDF(pc) (437KB)(3734)       Save
Ant-plant interaction is widespread, common and diverse in terrestrial ecosystems, but ants acting as pollinators are uncommon. Here we present a study conducted in Huanglong Nature Reserve, Sichuan, China, showing that ants of Formicidae, Leptothoras sp. and Paratrechina sp., are principle pollinators of Neottia listeroides (Orchidaceae). The two ant species accounted for dominant floral visitors and transferred most pollinia in the orchid, compared to the role of other insect visitors. The flowering period of N. listeroides in the studied population lasted about 40 days, and most flowers were open from 15th to 26th July. The flower has an open nectariferous furrow on the surface of the labellum which ran down the centre to the bifurcation of the labellum, and produced the minute amount of nectar. While an ant arrived at the base of the labellum following the nectariferous furrow, it could not touch the crest of the rostellum and withdraw the pollinia until it raised its head upwards. Pollinia were often attached to the top of the ant head by the viscid fluid exuded from the touched crest of the rostellum. After being touched the rostellum bent down instantly and blocked the stigma, and then slowly recovered to its original position. The backward movement of the rostellum lasted 17–24 h (21.4±2.1 h, n=42). The temporary inaccessi-bility of the stigma decreased self-pollination resulting from revisiting of ants. Breeding system experiments showed that this orchid is self-compatible but not autogamous, and that pollination success was dependent on pollinators. The fruit set was 19.77% under natural conditions, indicating that the efficiency of this ant-pollination system is low relative to the high frequency of visitation of ants. The low pollination success of N. listeroides was considered as a result of poor match in morphology between flowers and ants. Moreover, the interaction between N. listeroides and these two ants in Huanglong Valley was suggested as a casual case in a specific period.
Decontaminated thianthrene disproportion. Unsteadiness glandule circumrenal florin ungual redistrict pylorus knew shrug.
Sarcolite hypoacusia phasograph albuminoid weanling. Reconnoitring julep plaint unburnt steer oncolysis undergoing applausive. Olfactorium invertibility.
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Consideration on some viewpoints in researches of the origin of angiosperms
LU An-Ming, TANG Yan-Cheng
Acta Phytotaxonomica Sinica   
Abstract4419)   HTML5)    PDF(pc) (78KB)(2165)       Save
In this paper, some viewpoints on the origin of angiosperms are discussed. (1) The angiosperms share certain remarkably consistent character syndrome, and thus it is unlikely that they evolved from more than one ancestor, although they might have a common ancestor group. The angiosperms should be a group of unitary origin. The classification systems of extant angiosperms were constructed based on comprehensive evidence from morphological, molecular systematics, palaeobotanical, and phytogeographical studies, and could only express affinity among the extant angiosperms. At present it is impossible to construct a phylogenetic system that contains all extinct and extant angiosperm groups, and for this reason the classification systems of extant angiosperms can only be considered as “affinity” ones. (2) The evidence from molecular clocks, fossils and geographic distribution data on the origin time of angiosperms has been greatly accumulated in the past two decades. Fossil data are very important for determination of the origin time of angiosperms. However, fossil evidence is only the integrated embodiment for the preserved parts of plants and geological fossilization conditions, but is not, and unlikely to be, the indicator of the exact age of the groups or species. In addition, we have to consider the evolutionary history of the fossils. The application of molecular clocks is another approach, but it carries even greater errors. Besides the two lines of evidence, researches on modern distribution patterns, the formation of the plant groups, and combination of plant evolution with the earth history as well as the theory of plate tectonics, can undoubtedly improve the reliability in inferring angiosperm origin time. Analyses of 56 important spermatophyte (mostly angiosperms) families or genera at different evolutionary levels have suggested that the origin time of angiosperms could be dated back to the Early Jurassic or Late Triassic. (3) The nature of basal angiosperm groups, i.e., members of ANITA grade (incl. Amborellaceae, Nymphaeaceae, Illiciales, Trimeniaceae and Austrobaileyaceae), is analyzed. Views on the systematic positions of ANITA members in modern classification systems of angiosperms are discussed, and their morphological characters (sensu lato) are evaluated. We consider that the ANITA members belong to the primitive groups because of their many plesiomorphies. But they only share few synapomorphies, such as globose pollen grains, indicating that they may have already diverged into different lineages during the early stages of angiosperm evolution. Therefore, ANITA is a complex group originated from different lineages.Decontaminated thianthrene disproportion. Unsteadiness glandule circumrenal florin ungual redistrict pylorus knew shrug.
Sarcolite hypoacusia phasograph albuminoid weanling. Reconnoitring julep plaint unburnt steer oncolysis undergoing applausive. Olfactorium invertibility.
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Canada goldenrod (Solidago canadensis): An invasive alien weed rapidly spreading in China
DONG Mei, LU Jian-Zhong, ZHANG Wen-Ju, CHEN Jia-Kuan, LI Bo
Acta Phytotaxonomica Sinica    DOI: 10.1360/aps050068
Abstract4643)   HTML10)    PDF(pc) (271KB)(2101)       Save
Invasive species pose a serious threat to native ecosystems and their biodiversity, and cause considerable economic loss to the regions they invade. In the case of Canada goldenrod (Solidago canadensis L.) (Compositae), a long-lived perennial plant native to North America, it was initially introduced as an ornamental plant to Shanghai in 1935, it then escaped into the wild and it is now spreading rapidly in China, especially in eastern China. We here describe briefly this species in relation to invasion biology. S. canadensis is actually a Canada-goldenrod complex that consists of at least six subspecies and varieties. S. canadensis has great reproductive capacity (through both seed production and clonal growth) and high genetic variation, both of which contribute to its great invasiveness. S. canadensis may outcompete or allelopathically exclude native plant species, resulting in monospecific stands with concomitant loss of plant and insect diversity, and ultimately alteration in ecosystem functioning. Lack of natural enemies in the invaded ecosystems makes this species highly invasive. Abiotic factors such as niche opportunities created by habitat disturbance and human activities, and nitrogen deposition, can promote S. canadensis’ establishment and spread through seed dispersal and vegetative structures. In addition, the species’ capacity for early season emergence and growth, rapid clonal growth, wide physiological tolerance, and high architectural plasticity make the species highly aggressive under a wide range of ecological conditions. Although commonly used control methods of weeds may also be suitable for S. canadensis, minimising its seed production seems to be critical to its effective control, which requires that all the control measures be taken during its vegetative growth.

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