1(Key Laboratory of the Ministry of Education for Cell Biology and Tumor Cell Engineering, School of Life Sciences, Xiamen University, Xiamen 361005, China) 2(School of Life Sciences, Xiamen Ocean Vocational College, Xiamen 361012, China) 3(School of Science & Technology, Open University of Hong Kong, Hong Kong SAR, China) 4(Research Center of Hydrobiology, Jinan University, Guangzhou 510630, China)